Markus arrives, fights Michael and impales him. Selene and Michael visit Macaro, who refuses to assist Selene in killing his sons. After Selene and Michael leave, Markus arrives and drinks Tanis' blood to learn Selene and Michael's location, killing Tanis. Viktor killed Selene's family because they knew the prison's location, but made Selene into a vampiress with the location of the prison encoded in her blood. Tanis further reveals that Selene's father was the architect who built William's prison and that the pendant is the key. Due to William's destructiveness, Markus approached Viktor, then a dying mortal warlord, and offered to turn him and his army into vampires in exchange for tracking down and stopping William, and in destroying those he had infected. The first werewolves created by William were entirely animal and unable to return to their human forms. The third son remained human and gave rise to a line of mortal descendants including Michael, who turned into the first Lycan-Vampire hybrid. Markus was bitten by a bat and metamorphosed into a vampire, while William was bitten by a wolf and became a werewolf. Tanis reveals that Markus was the first vampire, one of the three sons of Macaro, the first immortal. Wanting to know why Markus wants it, the two travel to the hideout of the exiled vampire historian Andreas Tanis That night, Selene recalls seeing the pendant as a child, but does not know its significance. Using the knowledge obtained from Kraven's blood, Markus tracks the pair down and attacks them, but they escape and hide in a warehouse. Meanwhile, vampiress Selene takes Michael to a safe house. Micheal obtained the other half of the pendant after the death of Lucian.
Macaro examines Viktor's corpse and finds a metal disc inside it is the match to a pendant originally worn by Sonja. Alexander Corvinus, A.K.A Lorenz Macaro, the forefather to both the vampire and Lycan races, sends in a team of "cleaners" to investigate the aftermath from the battle in the Lycans' lair. In the present day, Markus awakens and kills regent Kraven.
Despite Markus's defiance, Viktor orders that William be imprisoned in a secret location forever. In 1202, an army led by the three vampire elders ( Markus, Viktor, and Amelia) arrives at a village full of Werewolves and capture their target: Markus's twin brother, William Corvinus, the first and most powerful werewolf.
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